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FHL 101 Introduction to Marine Biology in the San Juan Islands (1) NSc
One-weekend, one-credit course designed to introduce students to the marine ecosystem of the San Juan Archipelago and the facilities of the Friday Harbor Laboratories, while doing basic exploration of marine biology. Students will spend two full-schedule days doing a variety of field and lab studies. Offered: Sp.

FHL 250 Marine Biology (5) NSc
Physical, biological, and social aspects of the marine environments. Topics include oceanography, ecology, physiology, behavior, conservation, fisheries, exploration, and activism. Field and laboratory exercises expose students to common marine biological techniques and the diverse marine ecosystems of the San Juan Islands, WA, and to service-learning with local schools. Course equivalent to: FISH 250/BIOL 250/OCEAN 250 (when 5 credits) . Course overlaps with: FISH 250/BIOL 250/OCEAN 250 (when 3 credits). Recommended: at least one quarter of introductory biology (more preferable). Offered: Sp.

FHL 275 Natural History of the Salish Sea (5) NSc
A field-oriented introduction to the natural history of the Salish Sea focused on marine habitats and species of cultural and ecological importance. Students live at the Friday Harbor Labs, allowing immersive learning on the shores of the San Juan Islands. Lectures, field trips and lab exercises develop observational skills and knowledge of coastal organisms and their relationships with the environment. Human impacts are also discussed. Offered: Sp.

FHL 305 Biology of Fishes (5) NSc
Covers the morphological, physiological, behavioral, and ecological diversity of fishes worldwide; provides a basic foundation for advanced courses in all areas of aquatic sciences. Participants engage in field-based collecting, copious dissection/illustration laboratories, and delve into scientific literature on this fascinating group's biology. Course equivalent to: FISH 311/BIOL 311. Recommended: Recommended: 10 credits biological science. Offered: ASp.

FHL 333 Science Writing for Diverse Audiences (3/5)
Examines the effective communication of scientific ideas in writing for both scientific and public audiences. Students read, analyze, and write diverse materials, learning to identify and practice concise, effective communication about marine science for a variety of audiences. The close-knit, hands-on FHL community creates an environment where students work together both inside and outside of class to build their skills. Course overlaps with: MARBIO 305. Offered: ASp.

FHL 350 Introduction to Research in Marine Biology (6) NSc
Learn skills important to research scientists while participating in a mentored, hands-on project. Includes gathering, manipulating, and presenting data, plus group discussions on research ethics and the role of research in career options and graduate programs. Focuses on research activities in collaboration with a mentor. Research fields and topics vary. Recommended: any 100- or 200-level laboratory-based science course, ideally basic biology or marine biology. Offered: ASp.

FHL 375 Marine Mammals of the Salish Sea (5) NSc
An overview of the marine mammals found in the Salish Sea: their biology, habitats and conservation. Field trips throughout the San Juan Archipelago allow observation of common species, involving identification and recording of behavior in nature. Course overlaps with: FISH 475. Prerequisite: BIOL 180, FISH 250, ENVIR 100, OCEAN 101, or OCEAN 200. Offered: Sp.

FHL 403 Novel Marine Ecosystems (5) NSc
Covers how novel marine ecosystems arise from extinctions, introductions, climate change, and habitat change. Students apply fundamental ecological concepts to investigate novel ecosystem dynamics and management. Uses place-based, field-based, and project-based learning. Teaches hands-on scientific and professional skills in ecological theory and practice. Prerequisite: either FISH 310, FISH 312, FISH 323, FISH 370/MARBIO 370/OCEAN 370, BIOL 356, OCEAN 320, or OCEAN 330; recommended: background in marine ecology; evolution; and ecosystems and organisms; and a course in quantitative reasoning at the level of either Q SCI 381 or STAT 311. Offered: ASp.

FHL 420 Marine Ecology of the Salish Sea (5) NSc
Field-oriented study of ecology of the Salish Sea. Students live and study at the Friday Harbor Laboratories, a seaside setting for immersive learning. Students observe relationships between coastal organisms and their environment, and explore processes by which these relationships change over time and location. Lecture, field, and laboratory activities focus on habitats and species of ecological, economic, and/or cultural importance. Prerequisite: FISH 250/OCEAN 250/BIOL 250 or BIOL 356. Offered: jointly with BIOL 410; Sp.

FHL 430 Marine Zoology (5) NSc
Survey of groups of invertebrate animals represented in the San Juan Archipelago; natural history, functional morphology, ecology, distribution, habitat, adaptation, trophic interrelationships, and evolution. Course overlaps with: FISH 310. Prerequisite: BIOL 445, which must be taken concurrently; permission of Friday Harbor director. Offered: jointly with BIOL 430.

FHL 432 Marine Invertebrate Zoology (9) NSc
Comparative morphology and biology of marine invertebrates with emphasis on field and laboratory studies. Representatives of all major and most minor phyla are collected, observed live, and studied in detail. Taken at Friday Harbor Laboratories. Cannot be taken if credit received for BIOL 434. Course overlaps with: FISH 310. Offered: jointly with BIOL 432; S.

FHL 435 Research Apprenticeship in Marine Science (15) NSc
Immersive quarter in research n close collaboration with one-three faculty mentors on a specific topic that varies with the apprenticeship. Students engage in laboratory or field research in marine science, involving gathering, analyzing, and communicating results as part of a research team. Offered: jointly with BIOL 435/MARBIO 435; ASpS.

FHL 440 Marine Botany (5) NSc
Survey of plants represented in marine environments; natural history; ecology, distribution, habitat, adaptation, and trophic interrelationships. Taken at Friday Harbor laboratories. Prerequisite: either BIOL 220, B BIO 220, or TESC 140; BIOL 430, which must be taken concurrently. Offered: jointly with BIOL 445.

FHL 446 Marine Botany: Diversity and Ecology (9) NSc
Introduces marine plants - the diversity, life histories, physiology, and ecology of seaweeds, seagrasses, and marsh plants in the Salish Sea. Emphasizes field and laboratory work, including the application of molecular techniques to explore algal diversity and evolution. Offered: S.

FHL 450 Nearshore Ecology Research Experience (6) NSc
A hands-on introduction of "doing science" in close collaboration with a faculty member. Students engage in research around a thematic focus of ecology of nearshore marine ecosystems, with lab experiments and field work. Students gather, analyze, write up, and present their own data. Offered: jointly with MARBIO 450; ASp.

FHL 460 Aquatic Animals Research Experience (6) NSc
A hands-on introduction of "doing science" in close collaboration with a faculty member. Students engage in research around a thematic focus of aquatic animals, with lab experiments and field work. Students gather, analyze, write up, and present their own data. Offered: jointly with MARBIO 460; ASp.

FHL 468 Advanced Topics in Ecology and Biomechanics (9, max. 27)
Focuses on various aspects of ecology and biomechanics of marine organisms, such as experimental ecology, ocean acidification, deep sea biodiversity, or biomechanics. Taught at Friday Harbor Laboratories. Offered: S.

FHL 469 Ecology and Human Interactions in the Salish Sea (5) NSc
Explores complex, interrelated processes of marine social-ecological systems using the Salish Sea as a case study. Emphasis on building knowledge of ecological processes, familiarity with primary literature, and a sense of place in Salish Sea ecosystems through hands-on field activities and diverse course content. Prerequisite: either FISH 270/OCEAN 270/MARBIO 270, BIOL 220, B BIO 220, or T BIOL 140; recommended: basic biology and ecology coursework. Offered: jointly with MARBIO 469; A.

FHL 470 Research in Marine Biology (1-15, max. 15) NSc
A hands-on introduction of "doing science" in close collaboration with a faculty member. Students engage in research in marine biology; credits vary with the scale of the project with may include lab experiments and field work. Students gather, analyze, write up, and present their own data. Offered: jointly with MARBIO 470; ASp.

FHL 471 Comparative Anatomy and Physiology of Marine Organisms (5) NSc
Challenges and opportunities presented by the marine environment. Relationship between organismal form and physiological function in marine animals and plants. Field trips for collection of live local organisms. Physiology experiments and CT scanning. Offered at Friday Harbor Laboratories. Course overlaps with: BIOL 467 and FISH 324. Prerequisite: either BIOL 220, BIOL 240, or FISH 270/MARBIO 270/OCEAN 270; recommended: Q SCI 381 or equivalent. Offered: jointly with BIOL 471; A.

FHL 472 Friday Harbor Labs Research Apprenticeship (15, max. 30) NSc
A full-time, quarter-long program where apprentices work intensively on learning methods and carrying out hands-on research around one theme. Themes vary with the quarter and involve either extensive field or laboratory work or both. Students are trained in context, tools, data-gathering, and creation of research products appropriate to the theme. Recommended: strong background in core biological sciences. Offered: AWSp.

FHL 490 Marine Sciences Seminar (1, max. 3) NSc
Students attend seminars by marine scientists each week and ask questions about their seminars. Offered: ASpS.

FHL 492 Ecology and Conservation of Marine Birds and Mammals (9/15) NSc
An intensive, field-based course that offers motivated students the opportunity to learn about these ecologically and culturally important marine animals and the conservation problems they face. Emphasizes first-hand learning and individual research projects. Offered: jointly with FISH 492/MARBIO 492; S.

FHL 495 Special Topics in Natural Science (3-5, max. 10)
Special topics courses in the natural sciences that take advantage of unique opportunities at the Friday Harbor Laboratories. Offered: A.

FHL 528 Advanced Topics in Fish Biology (9, max. 27)
Focuses on various aspects of fish biology, such as functional morphology, sensory biology, or swimming mechanics. Taught at Friday Harbor Laboratories. Offered: S.

FHL 536 Comparative Invertebrate Embryology (9)
Studies diversity in developmental patterns in major marine taxa. Analyzes evolutionary changes in development, with emphasis on observation of live embryos and larvae. Offered: jointly with BIOL 536; S.

FHL 548 Advanced Topics in Evolution and Development (9, max. 27)
Focuses on various aspects of evolution and development of marine organisms, such as larval biology or evolution and development of metazoans. Students work on individual projects during the course. Taught at Friday Harbor Laboratories. Offered: S.

FHL 568 Advanced Topics in Ecology (9)
Introduces various aspects of marine ecology, such as experimental ecology, ocean acidification, and disease ecology. Taught at Friday Harbor Laboratories. Topics may change from year to year but will always focus on advanced topics in ecology. Recommended: college-level background in biology and ecology. Offered: S.

FHL 578 Special Topics in Science Communication (9) NSc
This FHL course focuses on different aspects of linkages between scientists and the public, including multiple ways to communicate science. Offered at Friday Harbor Laboratories; see FHL website for topic for next summer.

FHL 585 Advanced Topics in Neurobiology and Physiology (9, max. 27)
Focuses on various aspects of neurobiology or physiology of marine organisms, such as neurobiology, ethology, or marine chemosensory biology. Taught at Friday Harbor Laboratories. Offered: S.